"Anillini are a tribe of minute ((2.2-1.4mm) euedaphic ground beetles (Carabidae) characterized by the loss of eyes, loss of wings and high levels of local endemism. Despite their presumed low dispersal, they have a nearly cosmopolitan distribution, including isolated islands such as New Zealand and New Caledonia. The monophyly of Anillini was well-supported, and its age was consistent with a Gondwanian origin of the main lineages and an initial diversification at ca. 100Ma representing the split between the eyed Nesamblyops (New Zealand) and the remaining Anillini. The subsequent diversification, including the split of the Nearctic Anillinus and the subsequent splits of Palaearctic lineages, was dated to between 80 and 100Ma and thus was also compatible with a tectonic vicariant origin. On the contrary, the estimated age of the New Caledonian blind Orthotyphlus at ca. 30±20Ma was incompatible with a vicariant origin, suggesting the possibility of trans-oceanic dispersal in these endogean beetles. Specimens were generally collected from soil or forest litter."

Carmelo Andújar, Arnaud Faille , Sergio Pérez-González , Juan P. Zaballos , Alfried P. Vogler, Ignacio Ribera

in "Gondwanian relicts and oceanic dispersal in a cosmopolitan radiationof euedaphic ground beetles" ,Molecular Phylogenetics and Evolution 99 (2016) 235–246.