Référence

 Kazuaki Ohashi,Tatsuki ikeda, Hiroyoshi Ohashi, Tomoyuki NemOtO, Hirokazu muragaki, Haruna izumi, Haruna kObayashi, Koji Nata, Natsumi satO and Maika suzuki. 2018. Phylogenetic Analyses for a New Classification of the Desmodium Group of Leguminosae Tribe Desmodieae. J. Jpn. Bot. 93(3): 165–189.